AUDL 4007 & GS12  – 2015

Auditory perception

UCL Speech, Hearing and Phonetic Sciences
Chandler House
2 Wakefield Street
London WC1N 1PF

This unit will provide advanced training in aspects of auditory perception that are particularly relevant to understanding both normal and impaired auditory function. An emphasis will be placed on the fundamental aspects of hearing such as frequency analysis, temporal analysis, pitch perception, intensity perception and binaural processing.
Course Structure: The course is run on Thursday afternoons in the second term only, starting on 15 January 2015. There is no meeting on 22 January, but there will be one on 19 February, even though it is reading week. The course comprises a mixture of lectures, demonstrations, laboratory sessions and tutorials. Unless otherwise noted, lectures will be given by Professor Stuart Rosen.

Venue and scheduling: Sessions start at 14:00 in Chandler House, taking place in Room G15 and then moving to Room B07, the basement teaching laboratory.

Week Date Lecture Topic Laboratory Course Work (CW)
1 15 January

Introduction to the course

The auditory periphery

Mathematical interlude: continuous spectra and adding levels

Review signals & systems Choose a paper
2 22 January No meeting
3 29 January Envelope & Temporal Fine Structure (TFS) A cochlear simulation Paper approved by 28 Jan
4 5 February

Frequency selectivity

Intensity & Loudness

Measuring frequency selectivity using notched noise Hand in CW 1 by noon on 6 February
5 12 February Binaural hearing (Prof D McAlpine) CW 1 returned with comments
6 19 February Temporal resolution Adaptive techniques using gap detection CW 1 handed in re-written
7 26 February Pitch perception (Prof A Faulkner)

Measuring fundamental frequency in various sounds Choose paper 2
8 5 March Psychoacoustics of hearing impairment

Paper 2 approved by now

9 12 March

Cochlear Implants

Listening to speech in noise with cochlear implant simulations

10 19 March Auditory Scene Analysis (Dr M Chait) Bring essay 2 for reading in lab
11 26 March Hearing speech in noise Listening to speech in different kinds of 'noise' CW 2 handed in re-written

Stuart Rosen 020 7679 4077 course organiser
Dave Cushing 020 7679 7400 laboratory technician

Main text: Plack C. (2014) The Sense of Hearing, 2nd edition. Erlbaum.
The first edition should be adequate if you can't obtain the newer one.

Supplementary Reading
  • BCJ Moore (ed) (1986) Frequency Selectivity in Hearing. London: Academic Press. Does what it says on the tin, with thorough accounts of the psychoacoustics of frequency selectivity, plus more.
  • S Gelfand (2004) Hearing: An Introduction to Psychological and Physiological Acoustics, Fourth Edition, Revised and Expanded. A nearly complete resource for anatomy, physiology and psychoacoustics. Especially good on the middle ear.
  • R Plomp (2002) The Intelligent Ear, Lawrence Erlbaum, Mahwah New Jersey. A very accessible account of hearing.
  • BCJ Moore (2003). An Introduction to the Psychology of Hearing, 5th ed. (Academic Press). A very complete guide to the literature, but at an advanced level.
  • WA Yost (2006) Fundamentals of Hearing: An Introduction, 5th ed. (Academic Press). A more elementary exposition. Particularly good on the anatomy & physiology.
  • J Schnupp, E Nelken & A King (2010) Auditory Neuroscience: Making Sense of Sound (MIT Press). A book with much more discussion of the neural substrates of perception, and focus on a more limited range of topics. Worth a look. There is also an extensive web site containing animations and demonstrations.
  • D Moore, P Fuchs, A Palmer, A Rees & C Plack (2010) Oxford Handbook of Auditory Science: The Ear, The Auditory Brain, Hearing

  • 2 pieces of coursework, each worth 15% of the final mark (max 1000 words each), both based on doing the work of a science journalist by summarising a chosen paper in a style suitable for a quality daily newspaper. You will received feedback on one version of this, and re-write.
  • A 2-hour written paper (70%)
You must pass the final exam to pass the course.

Final exam

The final exam will take place in the main exam period. It will consist of a set of 8 essay type questions, all of which you must answer. This allows 15 mins/question. A greater depth of understanding will be expected for students in AUDL GS12. Generally speaking, there is a straightforward question associated with one or more lecture topics, as for the exams in 2009 and 2008. The exam in 2007 was in a slightly different format. Other exams are available on the library web site. Use the code AUDL4007.

Here are some example questions with tips for model answers.

Course work

Write two articles for a lay audience summarizing the crucial results of a journal article relevant to this course, in a style appropriate for a quality daily newspaper The use of graphics and diagrams is more than welcome, but is not required. Below you will find a few articles that would be appropriate, but please choose your own. You must email the .pdf first so I can assess its suitability. A good source of relevant papers can be found in the following journals, although other,more general journals, can sometimes have appropriate studies too (e.g., Nature and Science). You can gain access to these through the UCL library page

  • Journal of the Acoustical Society of America in the Psychological Acoustics section. This is the pre-eminent journal in the field with a wide variety of studies concerning both normal and impaired hearing.
  • The Journal of Speech, Language and Hearing Research
  • Ear & Hearing Oriented more towards clinically-relevant issues.
  • International Journal of Audiology
  • JARO (Journal of the Association for Research in Otolaryngology)
  • Hearing Research
  • Audiology & Neuro-otology
  • Attention, Perception & Psychophysics (formerly known as Perception & Psychophysics )

There are also often papers related to auditory perception in more general journals like The Proceedings of the National Academy of Sciences, Current Biology, Nature, etc. The following are examples of articles that would be appropriate, but please do not use these. Get an article from within last 3 years or so. Ensure that the article you select is not a review article in which much of the work is done for you already! It must be a primary research paper, although an appropriate meta-analysis of a some clinical procedure might be OK. Read any article you propose to use beforehand to ensure that it can be explained to a lay audience! Note that you may need to read ahead of the syllabus for some topics. You may not choose a a paper which is relevant for your project, and the two papers must be in distinct areas.

  • Baker, R. J. & Rosen, S. (2002). Auditory filter nonlinearity in mild/moderate hearing impairment. Journal of the Acoustical Society of America, 111, 1330-1339.
  • Kong, Y. Y., Cruz, R., Jones, J. A., & Zeng, F. G. (2004).Music perception with temporal cues in acoustic and electric hearing. Ear and Hearing, 25, 173-185.
  • Musiek, F. E., Shinn, J. B., Jirsa, R., Bamiou, D. E., Baran, J. A., & Zaidan, E. (2005). GIN (Gaps-In-Noise) test performance in subjects with confirmed central auditory nervous system involvement. Ear and Hearing, 26, 608-618.
  • Neuman, A. C., Haravon, A., Sislian, N., & Waltzman, S. B. (2007). Sound-direction identification with bilateral cochlear implants. Ear and Hearing, 28, 73-82.
  • van Hoesel, R., Ramsden, R., & O'Driscoll, M. (2002). Sound-direction identification, interaural time delay discrimination, and speech intelligibility advantages in noise for a bilateral cochlear implant user. Ear and Hearing, 23, 137-149.
  • Yasin, I. & Plack, C. J. (2005). Psychophysical tuning curves at very high frequencies. Journal of the Acoustical Society of America, 118, 2498-2506.

Articles should be 1000 words or less. Your article should introduce the basic topic and discuss the motivation of the author(s) in doing the study, as well report the results and their implications. Ensure you use language any reasonably well-educated person could understand. Do not simply paraphrase (or outright plagiarise!) the author(s). Unlike an article in the popular press, you should have a reference list. An appropriate style can be found in this example, this one or this one, all from Nature News. Again, use the library page to access the ejournal and locate the full article by date.

You will submit your work in hard copy and in an email to me. Please also submit a hard copy of the article your essay was based on. The following deadlines are crucial:

  • Starting right now: Choose a paper and email the title and the pdf to for approval by 28 January. Use the subject header AUDL4007 coursework I or it may get lost in my InBox!
  • 5 February: Hand in essay 1 by the start of class
  • 12 February: Essay 1 returned with comments
  • 19 February: Hand in rewritten final version of essay 1 to be marked
  • 27 February: After looking at the comments on essay 1, choose and have approved a second paper (in a different topic area) by email to, using the subject header AUDL4007 coursework II
  • 20 March: Bring two hard copies (double-spaced) of essay 2 to the lab session, where fellow students will read them and give you feedback. Also please bring in one hard copy of the paper you are discussing.
  • 27 March: Hand in final version of essay 2 to be marked

Week by Week

This website will be updated frequently though the term. You can see how the teaching in previous years has run for 2014, 2013, 2012 and 2011. Journal articles that are suggested reading are readily accessible through the reading list in Moodle.

Week 1: Introduction to the course, a review of peripheral auditory physiology & technical preludes to frequency selectivity

Suggested readings
  • You need to be reasonably familiar with the basic concepts of signals & systems analysis, as detailed in Signals and Systems for Speech and Hearing. A lab session will review this material.
  • Chapters 1-3 in Plack also summarise the basic aspects of acoustics, signals & systems you need to be familiar with, with Chapter 4 covering the basic physiology.
  • Schnupp et al. too cover these areas well, with a different focus and more emphasis on the physiology of the ear.
  • Yost's chapters 6-9 on the anatomy and physiology are very clear, extensive and easy to get to grips with.
  • Chapter 12 in Signals and Systems for Speech and Hearing, 2nd edition gives a thorough summary of the ear as a signal processor.

Week 2: Envelope and Temporal Fine Structure (TFS)

Suggested readings
  • Rosen, S. (1992) Temporal information in speech: Acoustic, auditory and linguistic aspects. Philosophical Transactions of the Royal Society of London, Series B, 336: 367-373.

Week 3: Frequency selectivity & Intensity and loudness

Read Ch 5 & 6 in Plack.

Week 4: Binaural hearing (Prof D McAlpine)

Read Ch 9 in Plack. Ch 5 in Schnupp et al. could also be useful.

Week 5: Temporal resolution

Read Ch 8 in Plack.

Term ends 27 March 2015

Other links to material concerning relevant topics

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